____life began remains a puzzle to us横线上应该填什么?

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____life began remains a puzzle to us横线上应该填什么?
____life began remains a puzzle to us
横线上应该填什么?

____life began remains a puzzle to us横线上应该填什么?
在复合句中作主语的句子称为主语从句,从句作主语,谓语动词用单数;通常由从属连词that,whether,连接代词who,whose,what,which,whatever(无论什么),whoever(无论谁),whichever(无论哪个)或连接副词when,where,why,how等引导.
How life began remains a puzzle to us.
生活如何开始对我们来说还仍是一个困惑.
how life began 做主语