The board of directors ___shaking heads at the chairman's speech. 填is 还是are?为什么?

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The board of directors ___shaking heads at the chairman's speech. 填is 还是are?为什么?
The board of directors ___shaking heads at the chairman's speech. 填is 还是are?
为什么?

The board of directors ___shaking heads at the chairman's speech. 填is 还是are?为什么?
is 主语为board 而不是directors

is 主语是the board
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are
The board of directors 董事会 由许多成员组成,后面的heads也表示用复数